Ioannis Rallis

@article{pistevos_unpublished_2026,

title = {Unpublished Mediterranean and Black Sea records of marine alien, cryptogenic, and neonative species (III)},

author = {Jennifer Pistevos and Argyro Zenetos and Yolanda Stranga and Adrian Teaca and Tatiana Begun and Domen Trkov and Joachim Langeneck and Vasilis Gerakaris and Yiannis Issaris and Polytimi-Ioli Lardi and George Oikonomidis and Maria Salomidi and Konstantinos Tsirintanis and Periklis Kleitou and George Constantinou and Christina Michail and Michel Michel Bariche and Alan Deidun and Alessio Marrone and Michail Ragkousis and Grigorios Skouradakis and Fiona Nash and Evangelos Markos Papadimitriou Digenis and Marius Skolka and Kremena Stefanova and Ana Fortič and Maria Garcia and Thanos Dailianis and Ioannis Rallis and Julian Evans and Maria Altamirano and Maria Rousou and Víctor Orenes-Salazar and Andrea Bonifazi and Louis Hadjioannou and Maria Zois ND Zotou Vlantaridis and Ernesto Azzurro and Patrick Schembri and Francesco Tiralongo and Georgios Chatzigeorgiou and Manfredi Di Lorenzo and Maria Fernandez and Antonio Di Franco and Martina Orlando-Bonaca and Antonio Calò and Christina Malefaki and Jose Antonio García-Charton and Orestis Papadakis and Athanasios Nikolaou and Andromachi Gkoulia and Alfonso Ramos-Esplà and Anna Maria Mannino and Desirèe Dimichele and Simone Spinelli and Elisa Martínez and Andrea Tursi and Annalisa Capasso and Georgios Christidis and Michele Solca and Mouna Rifi and Radhouan El Zrelli and Ramla Bouhlel and Riccardo Scalera and Ada Bandi and Anna Reboa and Antonios Mazaris and Ibrahim Iziani and Iro Anastasopoulou and Konstantinos Kavakakis and Leonardo Scavino and Lorenzo Merotto and Maria Yankova and Monica Montefalcone and Neža Leban and Nicola Ungaro and Paolo Mariottini and Allison Perry and Petros Stavrou and Valerio Mazzella and Violin Raykov and Jasmine Ferrario and Caterina Stamouli and Apostolos Siapatis and Emmanouela Vernadou and Claudia Bommarito and Giacomo Milisenda and Emanuele Somma and Stefania Russo and Stelios Katsanevakis et.al},

url = {https://www.reabic.net/journals/bir/2026/3/BIR_2026_Pistevos_etal.pdf

/wp-content/uploads/2026/09/2026-Pistevos-BIR-36.pdf},

doi = {10.3391/bir.2026.15.3.01},

issn = {22421300},

year  = {2026},

date = {2026-09-18},

urldate = {2026-01-01},

journal = {BioInvasions Records},

volume = {15},

number = {3},

pages = {533–572},

keywords = {},

pubstate = {published},

tppubtype = {article}

}

@article{chatzigeorgiou_non-indigenous_2026,

title = {Non-Indigenous Species in Four Major Greek Marinas: A Closer View of Polychaeta},

author = {Georgios Chatzigeorgiou and Ioannis Rallis and Maria Lampa and Wanda Plaitis and Argyro Zenetos},

url = {https://www.mdpi.com/1424-2818/18/4/197

/wp-content/uploads/2026/04/2026-Chatzigeorgiou-diversity-14.pdf},

doi = {10.3390/d18040197},

issn = {1424-2818},

year  = {2026},

date = {2026-03-31},

urldate = {2026-03-01},

journal = {Diversity},

volume = {18},

number = {4},

pages = {197},

abstract = {Ports and recreational marinas are established hotspots for the introduction and secondary spread of non-indigenous species (NIS), mainly via vessel transport and biofouling on artificial substrates. In the AlienPorts project, we surveyed fouling and associated macrobenthic assemblages in four major Greek marinas across distinct marine regions—Rhodes (Levantine), Heraklion and Piraeus–Zea (Aegean), and Patra (Ionian)—from 2021 to 2022, applying two complementary protocols: (i) photoquadrat surveys and (ii) scraped assemblage sampling. The photoquadrats recorded twelve NIS and seven cryptogenic species, which were dominated by sessile fouling taxa, especially ascidians. The scraped samples revealed 106 polychaete and amphipod species, including 25 taxa (23 NIS, two cryptogenic). Six of the polychaetes are new NIS for the Mediterranean, and thirteen of the taxa are newly recorded as NIS in Greek waters. Heraklion had the highest NIS richness (18 NIS + two cryptogenic), while the other marinas hosted approximately ten NIS/cryptogenic taxa. In contrast, NIS often dominated the population densities, with Zea being the most impacted based on the NIS to native species ratios for both diversity and abundance. The two methods detected largely distinct biodiversity components and shared only one NIS, underscoring a strong complementarity. The multivariate analyses confirmed significant differences among marinas (ANOSIM, p < 0.01). These results support integrated, multi-method monitoring to enhance the early detection and impact assessment under MSFD Descriptor 2.},

keywords = {},

pubstate = {published},

tppubtype = {article}

}

@article{koumpaki_macrobenthic_2025,

title = {Macrobenthic communities of the continental shelf of Heraklion Bay (Crete, Greece): bathymetric distribution and temporal trends},

author = {Eleni Koumpaki and Maria Maidanou and Georgios Chatzigeorgiou and Ioannis Rallis and Dimitra Mavraki and Costas Dounas and Christos Arvanitidis and Panayota Koulouri},

url = {https://bdj.pensoft.net/article/174931/

/wp-content/uploads/2025/12/2025-Koumpaki-BDJournal-78.pdf},

doi = {10.3897/BDJ.13.e174931},

issn = {1314-2828, 1314-2836},

year  = {2025},

date = {2025-12-18},

urldate = {2025-12-01},

journal = {Biodiversity Data Journal},

volume = {13},

pages = {e174931},

abstract = {The Gulf of Heraklion is one of the most well-studied coastal marine ecosystems in the eastern Mediterranean. It is an oligotrophic area, exposed to wave action where muddy sediments prevail and its coastal zone is relatively unaffected by major riverine inputs. However, it faces pressures deriving from urbanisation, tourism, small-scale fisheries and climate change. Amongst its biological components, macrobenthos is an important component of soft-bottom habitats of the continental shelf and considered to be an indicator of environmental and human-induced disturbance. Nevertheless, long-term data are limited, thus restricting our understanding of their temporal trends. This is of particular importance for the Mediterranean Sea which is characterised as a hot spot of biodiversity and climate change impacts on its biota. 

 The macrοbenthic faunal communities were investigated in the continental shelf of Heraklion Bay within the framework of the European funded project entitled MARine Biodiversity and Ecosystem Functioning leading to Ecosystem Services (MARBEFES), with the objective to enhance the understanding of marine biodiversity, ecosystem functioning and the ecosystem services they provide. Additional studies, with the same objectives, were carried under national and local monitoring programmes. Samples were collected from a specific transect of stations, named H2 (10, 20, 30, 40, 50, 100 and 200 m depth), located in the wider marine area of the wastewater treatment plant of Heraklion City. The transect was sampled during three different sampling surveys and periods: June 2010, June 2015 and August 2024. 

 A total of 30,280 individuals were examined belonging to the taxonomic groups of Annelida, Crustacea, Mollusca, Echinodermata, Nemertea and varia. The identification of the key taxonomic classes (Polychaeta, Bivalvia, Gastropoda, Scaphopoda and Malacostraca) revealed 231 macrobenthic taxa. Out of the 231 taxa, only 30 were common amongst all the three sampling periods, while Polychaeta was the most abundant macrobenthic group in all sampling periods.},

keywords = {},

pubstate = {published},

tppubtype = {article}

}

@article{montesanto_new_2022,

title = {New records of rare species in the Mediterranean Sea (December 2022)},

author = {Federica Montesanto and Marco Albano and Dενιζ Ayas and Federico Betti and Gioele Capillo and Melih Ertan Çinar and Maria Corsini-Foka and Fabio Crocetta and Ertan Dağli and Claudio D’Iglio and Markos Digenis and Branko Dragičević and Sergio Famulari and Deniz Ergüden and Antonio Giova and Valentina Giussani and Razy Hoffman and Igor Isajlović and Lovrenc Lipej and Raquel López-Esclapez and Francesco Mastrototaro and Alessandra Moreni and Victor Orenes-Salazar and Panayotis Ovalis Panayotis Ovalis and Wanda Plaiti and Juan A Pujol and Lotfi Rabaoui and Ioannis Rallis and Manja Rogelja and Serena Savoca and Grigorios Skouradakis and Francesco Tiralongo and Margherita Toma and Domen Trkov and Nicolas Ubero-Pascal and Lamia Yacoubi and Ferhat Yalgin and Sercan Yapici and Leon L Zamuda},

url = {http://imbbc.hcmr.gr/wp-content/uploads/2022/12/2022-Montesanto-MMS-79.pdf

https://ejournals.epublishing.ekt.gr/index.php/hcmr-med-mar-sc/article/view/32369},

doi = {10.12681/mms.32369},

issn = {1791-6763, 1108-393X},

year  = {2022},

date = {2022-12-19},

urldate = {2022-12-20},

journal = {Mediterranean Marine Science},

volume = {23},

number = {4},

pages = {968--994},

abstract = {This Collective Article presents information on 20 taxa belonging to five (5) Phyla: Cnidaria (2), Mollusca (8), Arthropoda (4), Echinodermata (1) and Chordata (5) recorded from the Alboran Sea to the Levantine Sea. These new records were found in seven (7) different ecoregions as follows: Alboran Sea: new record of the rare football octopus Ocythoe tuberculata in the harbour of Algeciras (Spain); Western Mediterranean Sea: first record of the isopod Arcturinella deltensis in the Ligurian Sea, and the third in the whole Mediterranean Sea; third Mediterranean site for the Muricidae Coralliophila ahuiri along the Italian coasts (Alghero); first and westernmost record of the goby Corcyrogobius liechtensteini from the Iberian Peninsula; new record of the sea slug Glaucus atlanticus from the coast of the Iberian Peninsula; first and easternmost record of Holothuria (Vaneyothuria) lentiginosa lentiginosa along the Italian coasts; Tunisian Plateau/Gulf of Sidra: seventh record of the oceanic squid Thysanoteuthis rhombus in Tunisia, and first for the Gulf of Gabes. Ionian Sea: second occurrence of the Atlantic tripletail Lobotes surinamensis in the Ionian Sea and first record from the Strait of Messina area; new record of the rare sea elephant Pterotrachea coronata from the Strait of Messina; Adriatic Sea: first record of a facies of football ascidian Diazona violacea in the South Adriatic Sea (Tremiti Island, Italy); two records of the sevengill shark Heptranchias perlo in Croatian waters after more than 70 years of absence of documented records in this area; first known occurrence of the nudibranch Jalonus hyalinus in Slovenian waters and also the first known occurrence of this species in the Adriatic Sea; first record of the nudibranch Okenia elegans in Slovenian waters; Aegean Sea: First record of the parasitic isopod Nerocila milesensis along Greek coasts (Crete island) as well as the second record worldwide; additional record of the brachyuran Paragalene longicrura collected from the Saronikos Gulf (Greece); first record of the siphonophore Rhizophysa filiformis in Greek waters as well as the second record of this species in the eastern Mediterranean basin; new record of the rare and protected angelshark Squatina aculeata along Turkish coasts; Levantine Sea: first record of the marine amphipod Caprella andreae in the Levantine Mediterranean shore of Israel, based on both morphological and molecular data; first occurrence of the cephalopod Tremoctopus violaceus along Turkish coasts, which confirms its presence in the north-eastern coasts of Turkey; record of a bloom of the thermophilic jellyfish Pelagia noctiluca in the north Levantine Sea.},

keywords = {},

pubstate = {published},

tppubtype = {article}

}

@article{rallis_early_2022,

title = {Early Succession Patterns of Benthic Assemblages on Artificial Reefs in the Oligotrophic Eastern Mediterranean Basin},

author = {Ioannis Rallis and Giorgos Chatzigeorgiou and Marta Florido and Francisco Sedano and Avgi Procopiou and Melina Chertz-Bynichaki and Emmanouela Vernadou and Wanda Plaiti and Panayota Koulouri and Costas Dounas and Vasilis Gerovasileiou and Thanos Dailianis},

url = {http://imbbc.hcmr.gr/wp-content/uploads/2022/05/2022-Rallis-jmse-35.pdf

https://www.mdpi.com/2077-1312/10/5/620},

doi = {10.3390/jmse10050620},

issn = {2077-1312},

year  = {2022},

date = {2022-05-01},

urldate = {2022-05-09},

journal = {Journal of Marine Science and Engineering},

volume = {10},

number = {5},

pages = {620},

abstract = {The colonization of artificial structures by benthic organisms in the marine realm is known to be affected by the general trophic patterns of the biogeographical zone and the prevailing environmental traits at the local scale. The present work aims to present quantitative data on the early settlement progress of macrofaunal benthic assemblages developing on artificial reefs (ARs) deployed at the Underwater Biotechnological Park of Crete (UBPC) in the oligotrophic Eastern Mediterranean. Visual census and subsequent image analysis combined with scraped quadrats were used to describe the establishment of the communities and their development over three consecutive campaigns, spanning 5 years post-deployment. Macroalgae consistently dominated in terms of coverage, while sessile invertebrates displayed different patterns over the years. Polychaeta and Bryozoa were gradually replaced by Cnidaria, while Porifera and Mollusca displayed an increasing trend over the years. Motile benthos was mainly represented by Mollusca, while the abundance of Polychaeta increased in contrast to that of Crustacea. For both sessile and motile assemblages, significant differences were observed among the years. The results of this study indicate that ecological succession is still ongoing, and further improvement in the monitoring methodology can assist towards a more accurate assessment of the community composition in complex AR structures.},

keywords = {},

pubstate = {published},

tppubtype = {article}

}

@article{sedano_comparing_2019,

title = {Comparing sessile benthos on shallow artificial versus natural hard substrates in the Eastern Mediterranean Sea},

author = {Francisco Sedano and Marta Florido and Ioannis Rallis and Free Espinosa and Vasilis Gerovasileiou},

url = {https://ejournals.epublishing.ekt.gr/index.php/hcmr-med-mar-sc/article/view/17897},

doi = {10.12681/mms.17897},

issn = {1791-6763, 1108-393X},

year  = {2019},

date = {2019-05-01},

urldate = {2020-08-21},

journal = {Mediterranean Marine Science},

volume = {20},

number = {4},

pages = {688},

abstract = {Artificial structures cover a considerable part of the Mediterranean coasts. In the Aegean Sea, most studies related to artificial structures have focused in vagile fauna on harbors and marinas but little attention has been given to the sessile biota on coastal defense structures. The aim of this work was to describe for the first time the shallow sublittoral sessile benthos on coastal defense structures in Crete (Eastern Mediterranean Sea) in order to identify potential differences in comparison to natural rocky substrates, adopting both a taxonomic and functional (i.e. macroalgal structural complexity) approach. Three shallow (1-3 m) localities were studied in the north coast and three in the south coast of the island (six localities in total). At each locality, two types of hard substrate were selected: an artificial coastal defense structure (rip-rap) and the nearest natural rocky substrates. The percent cover of sessile taxa was calculated using random points counts over photoquadrats (20 x 20 cm). The structure of the assemblage differed between artificial and natural habitats. Values of Shannon-Wiener’s diversity index and number of taxa were higher in natural substrates. In addition, cover of arborescent macroalgae was lower on artificial substrates. In conclusion, rip-raps do not function as surrogates of natural hard substrates in the study area since their shallow subtidal assemblages differ in terms of community structure, diversity and functionality. The deficient performance of such artificial structures could be attributed to the combined effects of abiotic factors and biotic processes, including substrate nature and roughness as well as differential grazing pressure.},

keywords = {},

pubstate = {published},

tppubtype = {article}

}